8 Archaea 4 Bacteria Planctomycetes SBZP_4958 6 Bacteria AC1 B04R032 6 Bacteria Planctomycetes vadinHA49 PHOS 22 Bacteria Gemmatimonadetes Gemm 7 Bacteria Proteobacteria Deltaproteobacteria Myxococcales 17 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae 40 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae 55 Bacteria Gemmatimonadetes Gemm 6 Bacteria WS3 PRR wb1_H11 7 Bacteria Gemmatimonadetes Gemm 10 Bacteria Gemmatimonadetes Gemm 19 Bacteria SAR406 AB16 noFP_H7 20 Bacteria Proteobacteria Deltaproteobacteria Myxococcales 7 Bacteria Gemmatimonadetes Gemm 7 Bacteria Firmicutes Clostridia Clostridiales Lachnospiraceae Coprococcus 10 Bacteria OP3 koll11 9 Bacteria Gemmatimonadetes Gemm 42 Bacteria Gemmatimonadetes Gemm 60 Bacteria Proteobacteria Deltaproteobacteria Desulfuromonadales Desulfuromonadaceae 16 Bacteria Proteobacteria Gammaproteobacteria 20 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 32 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 26 Bacteria Proteobacteria Deltaproteobacteria Desulfuromonadales 48 Bacteria Caldithrix Caldithrixae Caldithrixales BA059 33 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 60 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 5 Bacteria Proteobacteria Gammaproteobacteria Pseudomonadales Moraxellaceae Enhydrobacter 29 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobulbaceae 37 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobulbaceae 18 Bacteria Gemmatimonadetes Gemm 35 Bacteria Gemmatimonadetes Gemm 11 Bacteria OP3 PBS 9 Bacteria Fibrobacteres TG3 TG3 13 Bacteria Firmicutes Clostridia Clostridiales Peptococcaceae Pelotomaculum 6960 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Halomonadaceae 8 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 17 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae 18 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobulbaceae 12 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfovibrionaceae Desulfovibrio 86 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae Desulfovermiculus 24 Bacteria Fibrobacteres TG3 TG3 MAT 6 Bacteria Fibrobacteres TG3 TG3 15 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 26 Bacteria Gemmatimonadetes Gemm 7 Bacteria WS3 PRR GN03 20 Bacteria SBR1093 EC214 16 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae Desulfovermiculus 35 Bacteria Proteobacteria Gammaproteobacteria Chromatiales 20 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae Desulfovermiculus 19 Bacteria Proteobacteria Gammaproteobacteria 5 Bacteria Gemmatimonadetes Gemm 9 Bacteria Proteobacteria Deltaproteobacteria Bdellovibrionales Bacteriovoracaceae Bacteriovorax 9 Bacteria Proteobacteria Gammaproteobacteria Thiohalorhabdales Thiohalorhabdaceae Thiohalorhabdus 29 Bacteria Acidobacteria AT 12 Bacteria Gemmatimonadetes Gemm 9 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae 12 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae Desulfovermiculus 5 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 3 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 12 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae 9 Bacteria WS3 PRR GN03 14 Bacteria Proteobacteria Deltaproteobacteria Desulfuromonadales Desulfuromonadaceae 5 Bacteria SAR406 AB16 ST 10 Bacteria SAR406 AB16 ST 19 Bacteria Firmicutes Clostridia Halanaerobiales Halanaerobiaceae 9 Bacteria OP9 JS1 BA021 11 Bacteria Planctomycetes Planctomycetia Nitrospirales 6 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae 11 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae Desulfococcus 12 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Desulfobacteraceae 4 Bacteria Verrucomicrobia Verrucomicrobiae Verrucomicrobiales Verrucomicrobiaceae 17 Bacteria SAR406 AB16 ST 16 Bacteria Gemmatimonadetes Gemm 16 Bacteria OP1 KB1 5 Bacteria Fibrobacteres TG3 TG3 15 Bacteria GN04 GN15 4 Bacteria Proteobacteria Gammaproteobacteria 15 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales OM60 19 Bacteria Proteobacteria Deltaproteobacteria Desulfarculales Desulfarculaceae 7 Bacteria Proteobacteria Gammaproteobacteria HOC36 27 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales OM60 Congregibacter 29 Bacteria Caldithrix Caldithrixae Caldithrixales BA059 3 Bacteria Proteobacteria Deltaproteobacteria 8 Bacteria Proteobacteria Gammaproteobacteria 2086 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales Shewanellaceae Shewanella Shewanella_algae 6 Bacteria Spirochaetes Spirochaetes Spirochaetales Spirochaetaceae 28 Bacteria Proteobacteria Deltaproteobacteria Desulfarculales Desulfarculaceae 47 Bacteria Proteobacteria Deltaproteobacteria Desulfovibrionales Desulfohalobiaceae Desulfovermiculus 5 Bacteria Firmicutes Clostridia Thermoanaerobacterales Thermoanaerobacterales_Family_III 64 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 9 Bacteria Proteobacteria Gammaproteobacteria Chromatiales 9 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae 22 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 13 Bacteria Proteobacteria Deltaproteobacteria Desulfarculales Desulfarculaceae 4 Bacteria OP1 KB1 16 Bacteria Proteobacteria Gammaproteobacteria Legionellales Coxiellaceae 9 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales Alteromonadaceae Marinobacter 8 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Saccharospirillaceae 23 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Chromatiaceae Halochromatium 34 Bacteria Proteobacteria Gammaproteobacteria 9 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales OM60 65 Bacteria Proteobacteria Gammaproteobacteria Chromatiales 22 Bacteria Proteobacteria Deltaproteobacteria Desulfarculales Desulfarculaceae 4 Bacteria GN04 GN15 14 Bacteria Proteobacteria Gammaproteobacteria Chromatiales 21 Bacteria Proteobacteria Betaproteobacteria Burkholderiales Comamonadaceae Curvibacter 15 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Oceanospirillaceae 11 Bacteria Spirochaetes Spirochaetes Spirochaetales Spirochaetaceae Spirochaeta 113 Bacteria Proteobacteria Betaproteobacteria Burkholderiales Comamonadaceae 6 Bacteria Proteobacteria Zetaproteobacteria Mariprofundales Mariprofundaceae Mariprofundus 17 Bacteria Proteobacteria Gammaproteobacteria Chromatiales 5 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 22 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales Alteromonadaceae Alteromonas 12 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales OM60 11 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae Halorhodospira 19 Bacteria Firmicutes Clostridia Halanaerobiales Halanaerobiaceae Halanaerobium 7 Bacteria OP1 10 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales OM60 4 Bacteria Acidobacteria Sva0725 Sva0725 14 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales 6 Bacteria Chlorobi Ignavibacteria Ignavibacteriales 13 Bacteria Chlorobi Ignavibacteria Ignavibacteriales 7 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 6 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae 6 Bacteria Spirochaetes Spirochaetes Spirochaetales Spirochaetaceae Spirochaeta 7 Bacteria Gemmatimonadetes Gemm 16 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 11 Bacteria Proteobacteria Gammaproteobacteria HTCC2188 HTCC2089 11 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Halomonadaceae Halomonas 12 Bacteria Proteobacteria Gammaproteobacteria Xanthomonadales Sinobacteraceae 6 Bacteria Proteobacteria Deltaproteobacteria Clostridiales 6 Bacteria Spirochaetes Spirochaetes Spirochaetales Spirochaetaceae 13 Bacteria Proteobacteria Deltaproteobacteria Desulfuromonadales Pelobacteraceae Pelobacter 8 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae 10 Bacteria Proteobacteria Gammaproteobacteria Enterobacteriales Enterobacteriaceae 6 Bacteria Proteobacteria Gammaproteobacteria HTCC2188 9 Bacteria Proteobacteria Gammaproteobacteria Alteromonadales Alteromonadaceae 6 Bacteria Planctomycetes OM190 CL500 9 Bacteria Proteobacteria Deltaproteobacteria GMD14H09 7 Bacteria Proteobacteria Gammaproteobacteria Thiotrichales Piscirickettsiaceae 10 Bacteria Planctomycetes Planctomycetia Pirellulales Pirellulaceae 6 Bacteria Planctomycetes Phycisphaerae mle1 4 Bacteria Planctomycetes Planctomycetia Pirellulales Pirellulaceae 8 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Alcanivoracaceae Alcanivorax 7 Bacteria BRC1 PRR 3 Bacteria Fibrobacteres 29 Bacteria Bacteroidetes Saprospiraceae 4 Bacteria Bacteroidetes KSA1 8 Bacteria Bacteroidetes 27 Bacteria Bacteroidetes KSA1 7 Bacteria Bacteroidetes 6 Bacteria Bacteroidetes Saprospiraceae 4 Bacteria Bacteroidetes Saprospiraceae 12 Bacteria Bacteroidetes KSA1 5 Bacteria Bacteroidetes KSA1 22 Bacteria Bacteroidetes Chitinophagaceae 29 Bacteria Bacteroidetes Flavobacteriia Flavobacteriales Cryomorphaceae 4 Bacteria Bacteroidetes KSA1 12 Bacteria Bacteroidetes Saprospiraceae 6 Bacteria Bacteroidetes Saprospiraceae 31 Bacteria Bacteroidetes Flavobacteriia Flavobacteriales Flavobacteriaceae Robiginitalea 10 Bacteria Bacteroidetes Flavobacteriia Flavobacteriales Flavobacteriaceae Robiginitalea 7 Bacteria Bacteroidetes Flavobacteriia Flavobacteriales Flavobacteriaceae 10 Bacteria Verrucomicrobia Opitutae 8 Bacteria Bacteroidetes BME43 7 Bacteria Bacteroidetes Cytophagia Cytophagales Flammeovirgaceae 13 Bacteria Bacteroidetes Cytophagia Cytophagales Flammeovirgaceae 36 Bacteria Bacteroidetes Cytophagia Cytophagales Flammeovirgaceae 6 Bacteria Bacteroidetes Cytophagia Cytophagales Flammeovirgaceae 17 Bacteria Bacteroidetes Bacteroidia Bacteroidales 13 Bacteria Bacteroidetes Rhodothermaceae Salinibacter 15 Bacteria Bacteroidetes Rhodothermaceae Salisaeta 14 Bacteria Chlorobi OPB56 42 Bacteria Bacteroidetes Rhodothermaceae Salinibacter 6 Bacteria Bacteroidetes Rhodothermaceae Salisaeta 7 Bacteria Fibrobacteres Fibrobacteria Ucp1540 6 Bacteria Bacteroidetes Rhodothermaceae Salisaeta 20 Bacteria Proteobacteria Deltaproteobacteria 11 Bacteria Bacteroidetes Rhodothermaceae 10 Bacteria Elusimicrobia 18 Bacteria Bacteroidetes Rhodothermaceae Salisaeta 6 Bacteria Fibrobacteres Fibrobacteria Ucp1540 4 Bacteria OD1 8 Bacteria Actinobacteria Actinobacteria Actinomycetales Micrococcaceae Renibacterium 5 Bacteria GN04 GN15 9 Bacteria Planctomycetes Planctomycetia Gemmatales Gemmataceae Gemmata 10 Bacteria Chlorobi Ignavibacteria Ignavibacteriales OTUb7 6 Bacteria OP3 BD4 5 Bacteria OD1 ABY1 5 Bacteria Actinobacteria Actinobacteria Actinomycetales Corynebacteriaceae Corynebacterium 7 Bacteria Proteobacteria Deltaproteobacteria Myxococcales 6 Bacteria Actinobacteria Actinobacteria Actinomycetales Nocardiaceae Rhodococcus Rhodococcus_fascians 77 Bacteria Cyanobacteria Chloroplast Stramenopiles 16 Bacteria Actinobacteria Actinobacteria Actinomycetales Corynebacteriaceae Corynebacterium 28 Bacteria OD1 16 Bacteria Actinobacteria Actinobacteria Actinomycetales Propionibacteriaceae Propionibacterium Propionibacterium_acnes 5 Bacteria Lentisphaerae Lentisphaerales 5 Bacteria Cyanobacteria 4C0d MLE1 28 Bacteria Acidobacteria BPC102 B110 10 Bacteria OD1 ABY1 9 Bacteria OD1 ABY1 11 Bacteria Cyanobacteria Synechococcophycideae Pseudanabaenales Pseudanabaenaceae 22 Bacteria Cyanobacteria Synechococcophycideae Pseudanabaenales Pseudanabaenaceae Halomicronema 4 Bacteria Cyanobacteria Oscillatoriophycideae Oscillatoriales Phormidiaceae Microcoleus 6 Bacteria OP11 WCHB1 d153 8 Bacteria OD1 ABY1 5 Bacteria Firmicutes Clostridia Clostridiales Peptococcaceae Desulfitobacter 7 Bacteria Planctomycetes Phycisphaerae Phycisphaerales 18 Bacteria Cyanobacteria Synechococcophycideae Pseudanabaenales Pseudanabaenaceae Halomicronema 11 Bacteria Chloroflexi Anaerolineae OPB11 9 Bacteria Lentisphaerae Z20 6 Bacteria OD1 ABY1 4 Bacteria OP11 WCHB1 6 Bacteria Proteobacteria Deltaproteobacteria Myxococcales OM27 4 Bacteria Planctomycetes vadinHA49 DH61 36 Bacteria Chloroflexi Anaerolineae 26 Bacteria Actinobacteria Acidimicrobiia Acidimicrobiales 13 Bacteria Chloroflexi Anaerolineae 27 Bacteria Chloroflexi Anaerolineae 20 Bacteria 6 Bacteria Chloroflexi Anaerolineae OPB11 4 Bacteria Chloroflexi Anaerolineae 18 Bacteria Chloroflexi Anaerolineae GCA004 7 Bacteria Chloroflexi Anaerolineae 108 Bacteria Chloroflexi Anaerolineae GCA004 4 Bacteria Chloroflexi Anaerolineae 14 Bacteria Chloroflexi Anaerolineae SBR1031 A4b 7 Bacteria Planctomycetes Phycisphaerae Phycisphaerales 11 Bacteria Chloroflexi Anaerolineae OPB11 6 Bacteria Planctomycetes Phycisphaerae Phycisphaerales 26 Bacteria Chloroflexi Anaerolineae GCA004 8 Bacteria Planctomycetes Planctomycetia Pirellulales Pirellulaceae 10 Bacteria Actinobacteria Actinobacteria Actinomycetales 10 Bacteria Actinobacteria Acidimicrobiia Acidimicrobiales wb1_P06 5 Bacteria Actinobacteria Acidimicrobiia Acidimicrobiales C111 8 Bacteria Proteobacteria Deltaproteobacteria Desulfobacterales Nitrospinaceae Nitrospina 25 Bacteria Chloroflexi Anaerolineae GCA004 3 Bacteria Chloroflexi Anaerolineae 3 Bacteria Chloroflexi Anaerolineae OPB11 3 Bacteria Planctomycetes Phycisphaerae Phycisphaerales 4 Bacteria Proteobacteria Deltaproteobacteria Sva0853 JTB36 14 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae 20 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 10 Bacteria Nitrospirae Nitrospira Nitrospirales 4 Bacteria Proteobacteria Alphaproteobacteria 7 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae Rhodobaca 32 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 15 Bacteria Proteobacteria Alphaproteobacteria Rhizobiales Phyllobacteriaceae 6 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae 5 Bacteria Proteobacteria Gammaproteobacteria 7 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae 10 Bacteria Chloroflexi Anaerolineae OPB11 8 Bacteria Acidobacteria Solibacteres Solibacterales PAUC26f 17 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 7 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 3 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Hyphomonadaceae 6 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae Rhodovibrio 17 Bacteria Proteobacteria Epsilonproteobacteria Campylobacterales Helicobacteraceae 8 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae 32 Bacteria Proteobacteria Alphaproteobacteria Sphingomonadales Sphingomonadaceae Sphingomonas 26 Bacteria Chloroflexi Anaerolineae OPB11 12 Bacteria Chloroflexi Anaerolineae S0208 25 Bacteria Chloroflexi Anaerolineae OPB11 16 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 11 Bacteria Proteobacteria Deltaproteobacteria Myxococcales Syntrophaceae 14 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 68 Bacteria Proteobacteria Alphaproteobacteria Caulobacterales Caulobacteraceae 17 Bacteria Proteobacteria Alphaproteobacteria Kiloniellales 6 Bacteria OD1 ABY1 16 Bacteria Proteobacteria Alphaproteobacteria Rickettsiales Pelagibacteraceae 30 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 19 Bacteria Firmicutes Clostridia Clostridiales 31 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae Rhodovibrio 6 Bacteria Proteobacteria Alphaproteobacteria Rhizobiales Hyphomicrobiaceae Pedomicrobium Pedomicrobium_australicum 4 Bacteria Proteobacteria Alphaproteobacteria Kiloniellales 7 Bacteria Proteobacteria Alphaproteobacteria Rhizobiales Hyphomicrobiaceae 11 Bacteria Proteobacteria Alphaproteobacteria BD7 13 Bacteria Chloroflexi Anaerolineae Anaerolineales Anaerolinaceae T78 8 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae 11 Bacteria Proteobacteria Alphaproteobacteria Rhodospirillales Rhodospirillaceae Rhodovibrio 10 Bacteria Proteobacteria Alphaproteobacteria Sphingomonadales Erythrobacteraceae 3 Archaea Euryarchaeota Methanobacteria Methanobacteriales MSBL1 21 Bacteria Nitrospirae Nitrospira Nitrospirales 5 Bacteria Proteobacteria Gammaproteobacteria Chromatiales Ectothiorhodospiraceae 14 Bacteria Proteobacteria Alphaproteobacteria Rhodobacterales Rhodobacteraceae 10 Bacteria OP11 OP11 8 Archaea Euryarchaeota Halobacteria Halobacteriales Halobacteriaceae 4 Bacteria Planctomycetes Phycisphaerae Phycisphaerales Phycisphaeraceae 6 Bacteria Planctomycetes Phycisphaerae Phycisphaerales Phycisphaeraceae 4 Archaea Euryarchaeota Halobacteria Halobacteriales Halobacteriaceae Haloarcula 14 Bacteria Firmicutes Clostridia Clostridiales Anaerococcus 5 Archaea Euryarchaeota Halobacteria Halobacteriales Halobacteriaceae 14 Bacteria Proteobacteria Gammaproteobacteria Oceanospirillales Halomonadaceae 10 Archaea WCHD3 5 Archaea Euryarchaeota Methanomicrobia Methanosarcinales Methanosarcinaceae 5 Archaea Crenarchaeota MBGB 4 Archaea YLA114 6 Archaea Euryarchaeota Halobacteria Halobacteriales Halobacteriaceae 6 Archaea Euryarchaeota Methanobacteria Methanobacteriales WSA2 41 Archaea WCHD3 277 Bacteria Bacteroidetes Bacteroidia Bacteroidales Porphyromonadaceae Porphyromonas 8 Archaea Euryarchaeota Thermoplasmata E2 5 Bacteria Firmicutes Clostridia Clostridiales Ruminococcaceae 5 Archaea Euryarchaeota Thermoplasmata E2 11 Archaea WCHD3 24 Archaea WCHD3 10 Archaea WCHD3 4 Archaea Euryarchaeota DSEG ArcA07 6 Archaea Crenarchaeota MCG pGrfC26 23 Bacteria OD1 17 Bacteria Firmicutes Clostridia Clostridiales Clostridiaceae Clostridium